Shade leaves are often broader and thinner
Leaf size is one of their main adaptations. Broad and flattened leaves on plants such as umbrella leaf, hosta, and skunk cabbage function like huge solar collectors. Many leaves, such as those on ferns, aralias, and black cohosh, have dissected blades that are equally efficient, but also more wind-resistant. A large leaf divided into many small segments is less apt to be tattered or flattened in a summer storm. Leaves adapted to full sun are often smaller than shade-adapted leaves. Sun-adapted leaves have many layers of chlorophyll-rich cells, called palisade cells, piled one upon the other. This piggybacking is possible because the strong, direct sun can penetrate deep into the leaf. In this way, a small leaf can be very efficient. In contrast, shade-adapted leaves have a single layer of photosynthetic palisade cells, so the leaves must be proportionately bigger to accommodate the same number of cells and produce the same amount of food.
Leaves that are efficient in the sun are usually unable to function in the shade, and vice versa. For example, a potted ficus tree placed outside for its summer vacation gets lots of light, so it grows efficient, cell-packed leaves. In the autumn, when it's "back-to-school time," the ficus suffers a setback. Indoor light levels are low, and the high-efficiency leaves can't function, so the tree sheds them and produces new, larger leaves better adapted to less light.